dinosaurio animatronic fábrica de vestuario

Referral of the new-Fiberglass dinosaur-Dinosaur sculpture

Referral,of,the,new,specimen,to,Archaeopteryx,

The!taxonomy!of!Archaeopteryx!is!still!debated.!Almost!all!specimens!have,!at!

some!time,!been!described!as!separate!species,!and!several!even!as!different!

genera!(see!review!in!Wellnhofer1).!However,!it!is!generally!accepted!that!all!

avialan!theropods!from!the!Solnhofen!limestones!represent!members!of!the!

Archaeopterygidae,!and,!in!the!absence!of!a!detailed!analysis!of!the!taxonomy!of!

these!animals,!we!follow!the!recent!literature1–3!in!assuming!that!all!specimens!

represent!a!single!genus,!Archaeopteryx.!Unfortunately,!the!genus!Archaeopteryx!

(or!the!Archaeopterygidae,!in!cases!where!more!than!one!genus!has!been!

recognized)!has!not!been!satisfactorily!diagnosed!so!far,!with!the!latest!

diagnosis4!including!a!combination!of!characters,!none!of!which!is!unique!or!

apomorphic!for!this!taxon.!Differential!preservation!of!different!specimens!of!

Archaeopteryx!make!a!diagnosis!on!the!basis!of!shared!characters!of!all!

specimens!problematic,!but!there!are!several!characters!that!can!be!evaluated!in!

several!specimens!that!indicate!that!the!new!specimen!can!be!referred!to!this!

genus.!!

The!premaxillary!teeth!of!the!new!specimen!show!the!tooth!morphology!that!is!

typical!for!premaxillary!teeth!in!other!specimens!(London,!Berlin,!Eichstätt,!

Solnhofen,!and!Thermopolis!specimens),!in!that!the!teeth!have!a!straight!to!

slightly!bulbous!base!and!a!strongly!recurved!tip1,5!(Extended!Data!Figure!2b).!In!

nonWavian!theropods,!the!teeth!are!typically!more!gradually!curved,!whereas!

many!basal!avialans!have!also!gradually!curved!or!more!conical!teeth.!

In!the!jugal,!the!quadratojugal!process!is!short!and!very!slender!in!comparison!

with!the!suborbital!bar,!and!there!is!a!rounded!triangular!incision!between!the!

quadratojugal!process!and!the!postorbital!process,!forming!a!sharp!

anteroventral!margin!of!the!lower!infratemporal!fenestra!(Extended!Data!Figure!

2a).!This!morphology!is!also!seen!in!the!Eichstätt,!Munich!and!Thermopolis!

specimens2,6,7.!In!contrast,!in!nonWavian!theropods!and!basal!birds,!the!

quadratojugal!process!of!the!jugal!is!subequal!in!depth!to!the!suborbital!bar!or!

only!slightly!lower,!and!the!posterior!margin!of!the!postorbital!process!curves!

gently!into!the!dorsal!margin!of!the!quadratojugal!process.!Furthermore,!

Elzanowski4!noted!an!‘intermediate!process’,!a!small,!triangular!posterior!

projection!of!the!ischial!shaft,!as!a!diagnostic!character!of!the!Archaeopterygidae.!

Although!a!similar!process!is!present!in!Xiaotingia8,!it!is!especially!well!

developed!in!Archaeopteryx2,9–12,!and!this!character!is!also!present!in!the!new!

specimen!(Extended!Data!Figure!2c).!Finally,!asymmetric!tail!feathers,!as!found!

in!the!11th!specimen!(Fig.!2),!are!also!present!in!the!Berlin!specimen!(CF,!pers.!

obs.),!but!not!in!Anchiornis13.!Thus,!these!characters!strongly!indicate!that!the!

new!specimen!can!be!referred!to!the!Archaeopterygidae!and,!under!the!

assumption!of!a!single!genus,!to!Archaeopteryx.!The!species!taxonomy!of!

Archaeopteryx!is!in!need!of!revision1,2,14,!so!we!did!not!assign!the!new!specimen!

to!a!species.!



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